¹CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, Hubei, China
²Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, Wuhan 430074, Hubei, China
³Center of Economic Botany, Core Botanical Gardens, Chinese Academy of Sciences, Wuhan 430074, Hubei, China
⁴BTI Computational Biology Center, Boyce Thompson Institute, Ithaca, NY 14853, USA
⁵School of Integrative Plant Science, Section of Plant Biology and the L.H. Bailey Hortorium, Cornell University, Ithaca, NY 14853, USA
⁶Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumchi 830011, Xinjiang, China
⁷Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China
*Corresponding author. E-mail: sunhang@mail.kib.ac.cn,hcwang@wbgcas.cn
^†Both authors contributed equally to the study.