1Institute of Vegetables, Zhejiang Academy of Agricultural Sciences, Hangzhou 310021, China 2Institute of Vegetables, Ningbo Academy of Agricultural Sciences, Ningbo 315040, China 3State Key Laboratory for Quality and Safety of Agroproducts, Zhejiang Academy of Agricultural Sciences, Hangzhou 310021, China 4Present address: College of Life Sciences, China Jiliang University, Hangzhou 310018, China 5Biozeron Biotechnology Co., Ltd., Shanghai 201800, China *Corresponding author. E-mail: peixu@cjlu.edu.cn,yhwangsc@163. com