¹College of Eco-Environmental Engineering, Qinghai University, Xining 810000, China
²Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, Provincial Key Laboratory of Biotechnology of Shanxi Province, College of Life Sciences, Northwest University, Xi’an 710069, China
³College of Agriculture and Animal Husbandry, Qinghai University, Xining 810000, China
*Corresponding author. E-mail: jiaxianqing@nwu.edu.cn,1995990043@qhu.edu.cn
^†Both authors contributed equally to the study.