¹Haixia Institute of Science and Technology, State Key Laboratory of Agricultural and Forestry Biosecurity, Fujian Provincial Key Laboratory of Haixia Plant Systems Biology, Institute of Applied Ecology, Instrumental Analysis Center, Fujian Agriculture and Forestry University, Fuzhou 350002, China
²State Key Laboratory of Plant Environmental Resilience, College of Life Science, Zhejiang University, Hangzhou, Zhejiang 310058, China
³Department of Plant Physiology, Swammerdam Institute for Life Sciences, University of Amsterdam, Science Park 904, Amsterdam 1098 XH, The Netherlands
⁴Marine and Agricultural Biotechnology Laboratory, Institute of Oceanography, Minjiang University, Fuzhou 350108, China
⁵Shenzhen Institutes of Advanced Technology, Chinese Academy of Science, Shenzhen 518055, China
⁶Key Laboratory of Synthetic Biology, Center for Excellence in Molecular Plant Science, Institute of Plant Physiology and Ecology, Chinese Academy of Sciences, Shanghai 200032, China
⁷Department of Biological Sciences, Brock University, St. Catharines, ON L2S 3A1, Canada
⁸Laboratory of Southern Subtropical Plant Diversity, Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Sciences, Shenzhen 518004, China
⁹State Key Laboratory of Agricultural Genomics, BGI-Shenzhen, Shenzhen 518083, China
¹⁰Institute of Soil Water and Environmental Science, Volcani Institute, 68 HaMaccabim Road, PO Box 15159, Rishon LeZion 7505101, Israel
*Corresponding author. E-mail: yyliu@fafu.edu.cn
^†Minxuan Li and Sen Cai,Xu Bai contributed equally to the study.