¹Institute of Plant Resources, Key Laboratory of Biotechnology and Bioresources Utilization, Ministry of Education, Dalian Minzu University, Dalian, China
²State Key Laboratory of Tree Genetics and Breeding, National Engineering Research Center of Tree Breeding and Ecological Restoration, College of Biological Sciences and Technology, Beijing Forestry University, Beijing, China
³State Forestry and Grassland Administration Key Laboratory of Silviculture in Downstream Areas of the Yellow River, College of Forestry, Shandong Agricultural University, Taian, Shandong, Chin
⁴School of Biological Science and Technology, University of Jinan, Jinan, China
⁵Institute of Crop Germplasm Resources, Shandong Academy of Agricultural Sciences, Jinan, China
*Corresponding author. E-mail: ruan@dlnu.edu.cn
^†Zhi-Wei Wang,Ye Zhao and Peng Li contributed equally to the study.